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Structure of fish population usually varies
with ecological characteristics of the environment they
live in and also with genetic constitution of the
species. By the term “population structure”, we
mean data on size frequency distribution of a fish
population, weight distribution, sex and age
compositions, and length-weight relationship as
explained below:
Sex Composition: Sex composition of a balanced fish population is
generally 1:1. In other words, the ratio of male and
female members within total fish population is equal to
each other. If the size of fish population is lower
than the carrying capacity of their habitat, the female
ratio of sex composition is higher than the male ratio;
if it is greater than the carrying capacity then the
male ratio of sex composition is higher than the female
ratio. This is called as the “Cybernetic Population
Model” which in fact shows us that living organisms
have their own dynamic balance as long as there is no
human interference.
The male to female ratio of pearl mullet
population except for the reproduction period is around
1:1.85. In other words, 36% of the total fish
population is male whereas 64% is female. During the
reproduction period, sex composition changes in favor
of the male members. When sample fish groups are taken
from rivers as their primary habitats for reproduction
and analyzed further, it is seen that 64% of the total
fish population is male whereas 36% of it is female at
spawning. These ratios turn just the opposite of one
another during and beyond the reproduction season.
Since the pear mullet is not hermaphrodite in nature,
where does such a difference originate? Sampling errors
are those first come to mind. But various studies have
repeatedly provided similar results. In this case, we
cannot talk about any sampling errors. In fact, the
result of studies on spawning grounds of the pearl
mullet is also valid for a majority of migratory fish
species. Male members move to spawning grounds earlier
and leave them later than female members. So, studies
on spawning areas indicate that the male population is
more intensive in such places (Sarı, 1997). The end
results of studies on migratory fish species are also
supportive of this argument on the pearl mullet
population within which the sex ratio changes in favor
of male members, and in some cases it can even be
observed as 36:1 (Nikolskii 1980; Hofragle and Timmons
1989).
Age composition: Age
composition of fish populations varies with
reproduction, growth and mortality rates. As for the
maximum life span, it is more related with the genetic
constitution of fish.
There were various interpretations of maximum life span
until 1997 depending on techniques used or fish tissue
samplings taken for age determination of the pearl
mullet population. But according to a recent study
carried out by Sarı (1997), it is ideally best to use
otolith for age determination. It is further emphasized
that using scales or operculums does not provide
accurate results for age determination in most cases
due to the migratory nature of fish. After this study,
the maximum life span of pearl mullet has been
determined as a total of 7 years. Based on age
distribution of pearl mullet yield, the actual age
distribution was structured virtually using the virtual
population analysis technique. Results can be seen as
in the following diagram (Sarı, 2001).
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Length and weight distribution: Length
and weight distribution vary with genetic constitution
of the species, ecological characteristics of the
habitat, and population density available. In
nutritiously rich environments, if the size of
population is lower than the carrying capacity of it
then fish can grow faster, and therefore, their length
and weight composition will be higher than that of
those in nutritiously poor environments. Any changes in
such habitats in time will affect the length and weight
distribution of fish as well. For this reason, annual
length and weight distribution levels should be
observed on a regular basis. Just to give an idea about
annual length and weight distribution values, here
below let us refer to data provided by Sarı (1997). We do not give any further information about
these values here since detailed charts are already
provided at Growth
section,
Length-weight relationship: There
is a non-linear relationship between the length and
weight values of fish as given by the following
equation: W=a*Lb (W-weight of fish, L-length
of fish, a and b are regression constants). This means
that as the length of fish increases there will also be
an increase in the weight of fish according to the
exponential function of food capacity of the
environment and fish body shape. In the equation,
constant b gives the body shape of fish and
constant a gives the food capacity of the
environment.
The length-weight relationship of the pearl
mullet population has been calculated as W= 0.0912 L2.245
(Sarı, 1997). Parameters of this relationship can vary
with annual data, sampling size and sensitivity levels
of measurement and weighing.
Size of population, mortality and survival rates:
Size of pearl mullet population, mortality and survival
rates are determined by “Virtual Population Analysis”
technique which has been widely used in recent years.
After identifying size, age and growth characteristics
of fish mostly caught by professional fishery, these
values have been analyzed using the technique mentioned
above for the first time to determine the stock size of
total inland fish in our country.
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Ages
|
Fishing
(Number)
|
Stock Size
|
F
|
M
|
Z
|
S
|
| Number
|
(Ton) |
|
I
|
0
|
465393216
|
7581,256
|
0
|
0.2
|
0.2
|
0,819
|
|
II
|
70686168
|
381031744
|
14239,156
|
0.228
|
0.2
|
0.428
|
0.652
|
|
III
|
137780752
|
248351168
|
12131,955
|
0.924
|
0.2
|
1.124
|
0.325
|
|
IV
|
36410304
|
80736520
|
4685,141
|
0.677
|
0.2
|
0.877
|
0.416
|
|
V
|
12132344
|
33575916
|
2199,223
|
0.503
|
0.2
|
0.703
|
0.495
|
|
VI
|
6524455
|
16620961
|
1228,455
|
0.561
|
0.2
|
0.761
|
0.467
|
|
VII
|
1280590
|
7768686
|
637,498
|
0.200
|
0.2
|
0.400
|
0.670
|
|
Total
|
264814624
|
1233478144
|
42702,684
|
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| Virtual
Population Analysis (VPA) on sampling from
commercial fishing data between 1994-1995 and 1995-1996
fishign season
(M-Natural mortality, F-Fishing mortality, Z-Total
martality (M+F), S-Survial ratio (S= exp(-Z) |
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